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Lightroom for organizing, basic/medium editing such as white balance and tone adjustments, PhotoShop for more advanced stuff such as merging panoramas, HDR's, merging multiple bracketed exposures as blended layers, etc.
I'd say since I started using Lightroom about 95% of my photo post-processing has been in Lightroom.
The latest version even does chromatic abberation removal, which used to be solely a PhotoShop domain.
So you can get away with Lightroom for most everything but not 100% of everything and eventually having Photoshop will be very useful.
Never used Aperture so can't comment on it compared to any other software.
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